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It is recommended to have the A/C on at all times to ensure that the gaskets do not dry out. It doesn't use much fuel anyway, and it keeps the car nice and dry inside, so you avoid the windows misting up.
The fan buttons are not directly controlling the fan speed with the A/C set to automatic. It controls the intensity of the system. At the lowest setting, it will run low fan speeds, and not try all that hard to keep the temperature correct. I usually prefer the second lowest intensity. Higher than that, and it becomes noisy and windy at times.
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